Jonathan Violin has served on our Board of Directors since January 2021. Dr. Violin is a co-founder of Quellis, and prior to the merger was its Treasurer and Secretary since December 2018. Dr. Violin has served as President and Chief Executive Officer of Viridian Therapeutics, Inc. (formerly Miragen Therapeutics, Inc.), a publicly traded biotechnology company, since January 2021, and was previously President and Chief Operating Officer of Viridian, from October 2020 until January 2021. Dr. Violin co-founded Viridian Therapeutics, Inc., a privately held biotechnology company (which was acquired by Miragen Therapeutics, Inc.) leading its operations since its founding in 2020. Dr. Violin also co-founded Dianthus Therapeutics, Inc., a biotechnology company, in May 2019, and co-founded and helped lead Trevana Inc., a biotechnology company, in various roles from 2008 until November 2018, most recently as Senior Vice President, Scientific Affairs and Investor Relations Officer. Dr. Violin holds a Ph.D. from the Department of Pharmacology at the University of California, San Diego, a Master of Business Administration with a concentration in Health Sector Management from the Fuqua School of Business, and a Bachelor of Sciences in Chemical Pharmacology from Duke University.