Dr. Langford previously served as Chief Scientific Officer at Rubicon Genomics and is its Co-Founder. John retired as a Professor at the University of Michigan from the Department of Molecular, Cellular, and Developmental Biology and was a leader in the use of electron microscopy and x-ray scattering to study chromosome fibers in living cells, and his models for nucleosome structure and the folding of nucleosomes into chromosome fibers were important milestones.